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Alpine black swallowtail

Papilio maackii

Lepidoptera / Papilionidae

The alpine black swallowtail has dark wings with green-blue reflective regions. Their reflected colour shifts with illumination angle, while chemical experiments show how Amur cork tree compounds stimulate female egg laying.

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Papilio maackii, dorsal view
Alpine black swallowtail studyPapilio maackii · Dorsal view.

At a glance

Family ↘
Papilionidae, the swallowtail family. [1]
Region ↘
East Asia; Taiwanese fauna treats its single historical record as accidental. [1]
Recorded host ↘
Phellodendron amurense, in a primary egg-laying study. [2]
Remarkable detail ↘
Reflected colour shifts toward shorter wavelengths under oblique illumination. [3]

Species identity

Papilio maackii, the alpine black swallowtail, belongs to Papilionidae. [1]

Green-blue reflective regions

Predominantly dark wings contain green and blue reflective regions. Their reflected hue varies with illumination angle, as measured in the wing-scale optical study. [1][3]

East Asian records

The Taiwanese fauna records the species in East Asia and classifies a single male collected in Taiwan in 1935 as an accidental occurrence. Japanese experimental material came from Honshu and Kyushu. [1][4]

Amur cork tree and other Rutaceae

In the host-selection study, females accepted Phellodendron amurense, the Amur cork tree, for egg laying. Larvae of both P. maackii and P. protenor developed well on this plant in the laboratory, although the latter species did not naturally use it in the study setting. [2]

Responses to leaf volatiles

In Japanese laboratory trials, tested females showed stronger host-searching and egg-laying responses to the three non-Citrus plants than to Citrus deliciosa. The tested alternatives were Phellodendron amurense, Zanthoxylum ailanthoides and Orixa japonica. [4]

Microscopic architecture and reflection

The green and blue wing regions have scales with microscopic ridges and cross-ribs. In the study, the reflected colour shifted toward shorter wavelengths as illumination changed from perpendicular to more oblique angles. The authors’ simplified chitin–air models linked differences in scale geometry to the green and blue reflection spectra. [3]

Species-specific responses to phellamurin

Phellamurin, a compound in Phellodendron amurense, stimulated egg laying in P. maackii but deterred P. protenor in the host-selection experiment. The same plant compound therefore had opposite effects on these closely related butterflies. [2]

Locating reflective areas

The green and blue regions reflect different portions of visible light. Changes in illumination can shift the apparent colour and brightness of these scaled surfaces. [3]

Reflective colour within dark wings

Green-blue flat colour regions preserve the reflective areas within dark wings. Bold contours define the silhouette and restrained internal lines retain the pattern boundaries. [1][3]

Protecting scaled wings

Protect scaled wing surfaces from handling and abrasion. Keep the specimen in a stable enclosure away from direct sunlight, heat and moisture. [5]

Host selection and optical mechanism

Why does the reflected colour change with angle? The optical study measured a shift toward shorter wavelengths under more oblique illumination and linked the reflection spectra to microscopic scale geometry. [3]

What stimulates egg laying on Amur cork tree? Phellamurin stimulated P. maackii females in the host-selection experiment, while deterring P. protenor. [2]

Chemical ecology and wing optics

The Taiwanese fauna records regional occurrence. Chemical-ecology studies examine host acceptance and plant volatiles, and the optical paper reports wing-scale architecture and angular reflection. [1][2][4][3]

  1. Butterfly Fauna of Taiwan, Volume I: Papilionidae, second edition ↗

    Taiwan Forestry Bureau · Accessed 2026-10-09

  2. Modulation of oviposition response of Papilio maackii and P. protenor to Phellodendron amurense ↗

    Journal of Chemical Ecology · Accessed 2026-10-09

  3. 3D Modelling for Photonic Crystal Structure in Papilio maackii Wing Scales ↗

    Materials · Accessed 2026-10-09

  4. Behavioral and Electrophysiological Study on Eight Japanese Papilio Species with Five Hostplant Volatiles and Linalool ↗

    Insects · Accessed 2026-10-09

  5. Caring for natural history collections ↗

    Canadian Conservation Institute · Accessed 2026-10-09

  6. About Butterflies ↗

    Smithsonian Gardens · Accessed 2026-10-09

Artworks in the collection

Framed and digital artworks featuring this species. Current options and prices are listed on Etsy.

Related species

Sources checked 2026-10-09. Illustrations · Luminary Oddities.

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